PHILADELPHIA (Interstat) — Winston Watkins threw for 254 yards and rushed for 72 yards, accounting for two touchdowns, as William & Mary defeated Villanova 35-32 on Friday night at Villanova Stadium.

Watkins completed 19 of 33 passes for the Tribe, including a 68-yard touchdown to Carson Jenkins and a scoring strike to Kai Austin. He also ran for a 1-yard touchdown. William & Mary led 19-17 after three quarters and outscored Villanova 16-15 in the fourth.

Pat McQuaide completed 17 of 30 passes for 270 yards and a touchdown for the Wildcats. Isaiah Ragland rushed for 94 yards and two scores, Ja’briel Mace added 57 yards and a touchdown, and Jake Hilton had four catches for 132 yards.

Villanova’s defense was led by Omari Bursey’s 15 tackles, one sack and a quarterback hurry. Ratu Delano Navoti added two sacks. William & Mary got eight tackles from Harrison Johnson and six from Bryce Barnes.

The win lifted William & Mary to 1-0; Villanova dropped to 0-1.