TOWSON, Md. (Interstat) — Beau Brungard accounted for 377 yards of total offense and Andrew Lastovka kicked a 42-yard field goal as time expired to lift Youngstown State to a 31-28 victory over Towson in a non-conference football game Saturday at Johnny Unitas Stadium.

The Penguins (2-1) drove 40 yards in the final 37 seconds to set up Lastovka’s game-winning kick, capping a back-and-forth contest that saw the lead change three times in the second half.

Brungard, a quarterback from New Middletown, Ohio, completed 24 of 41 passes for 277 yards and a touchdown while also rushing 18 times for 100 yards and two more scores. His 1-yard plunge in the first quarter opened the scoring, and his 2-yard run with 6:21 remaining in the game put the Penguins ahead for good at 31-28.

Youngstown State built a 7-0 lead after the first quarter, but Towson (2-1) responded with 14 unanswered points in the second quarter. Al Wooten II scored on a 2-yard run and John Dunmore connected with Kemarrion Battles on a 33-yard touchdown pass to give the Tigers a 14-7 halftime advantage.

The Penguins reclaimed the lead in the third quarter. Brungard hit tight end Alex McDonald for a 2-yard touchdown pass, and after Towson answered with a 19-yard scoring pass from Andrew Indorf to Dunmore, Lastovka nailed a 27-yard field goal to give Youngstown State a 17-14 edge.

Wooten’s second touchdown run, a 1-yarder early in the fourth quarter, put Towson back in front 21-17 before Brungard’s final rushing touchdown and Lastovka’s late field goal sealed the win.

Indorf completed 18 of 29 passes for 238 yards and a touchdown for Towson, but was intercepted once by Youngstown State defensive back Justin Wimpye. Wooten led the Tigers’ ground attack with 48 yards and two touchdowns on 12 carries.

Max Tomczak was Brungard’s top target, hauling in seven receptions for 116 yards for the Penguins. Jaceon Doss had a big day receiving for Towson, catching three passes for 120 yards.

The loss snapped a two-game winning streak for Towson, which was a 3.5-point favorite according to sportsbooks. The combined score of 59 points went over the pregame over/under line of 52.5.

Youngstown State, which improved to 2-1, will open Missouri Valley Football Conference play at home against South Dakota State on Oct. 4. Towson, also 2-1, will travel to face Bryant next Saturday.